Rachelle Lefevre plays the renegade vampire Victoria in the film Twilight (2008), based on the novel of the same name by Stephenie Meyer. Directed by Catherine Hardwicke, the film was shot mostly in and around Portland, Oregon. Rachelle wrote an impassioned letter to the director, explaining her desire to work with the filmmaker. She described the essence of her character as "pure evil, pure instinct, pure malice, and very feline". After reading that the author used the word "feline" to describe her character's agility, Rachelle watched lion attacks on YouTube to separate the movements of her character from those of normal people. She also took trapeze classes in preparation for the wire work in the film. Rachelle spent hours working on her costumes for her character in the film, and described herself as "obsessed" with vampires after reading Bram Stoker's Dracula at the age of 14. She participated in a promotional tour in November 2008, where she met and signed autographs for Twilight fans at Hot Topic stores in the U.S..
Rachelle has at times been overwhelmed by the scrutiny she has received from her involvement with Twilight. "It's the closest I will ever come in my life to being a rock star," she told The Canadian Press, describing an appearance on MuchMusic, where over 1,500 fans showed up in Toronto. Overall though, she describes herself as delighted by the attention and excitement of the fans. Over 2,500 fans showed up at a Wal-Mart in Salt Lake City, where Lefevre appeared to promote the DVD release of the film.
Rachelle will appear in the film's sequel, based on Meyer's second novel, New Moon. She completed shooting the film in Vancouver in May 2009. Rachelle will not reprise her role of Victoria in Eclipse, the third film of the Twilight series, but will be replaced by Bryce Dallas Howard. Summit Entertainment, the studio behind the films, attributed the change to scheduling conflicts; Barney's Version and Eclipse both begin filming on 17 August 2009. Rachelle responded that she was "stunned" by the decision and "greatly saddened" not to continue her portrayal of Victoria, and never thought she would "lose the role over a 10 day overlap", in a statement to Access Hollywood. The studio responded in a statement, "The Twilight Saga: Eclipse is an ensemble production that has to accommodate the schedules of numerous actors while respecting the established creative vision of the filmmaker and most importantly the story." Rachelle appeared at the 2009 San Diego Comic-Con to promote New Moon the weekend before. Enthusiastic fans of the series reacted to the news with online petitions urging Rachelle's return, and "Bring Back Rachelle" became a top-trending topic on Twitter in the afternoon of 29 July 2009.
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